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For example, the tripeptide glutathione is synthesized in two steps from free amino acids, by two enzymes: glutamatecysteine ligase (forms an isopeptide bond, which is not a peptide bond) and glutathione synthetase (forms a peptide bond)

Neurobiol Dis (2012) 46(3):62534.10.1016/j.nbd.2012.02.012 22 ShatilloAKorolevaKGiniatullinaRNaumenkoNSlastnikovaAAAlievRRet alCortical spreading depression induces oxidative stress in the trigeminal nociceptive system