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Characterization of AUA decoding with modified mRNAs To examine the effect of the additional hydrogen bonding mediated by the ava 2 C side chain on AUA decoding, we performed an A-site tRNA-binding experiment using a series of synthetic mRNAs bearing 2-OH (control), 2-H, 2-OMe or 2-F at the residue 3-adjacent to the A-site codon (Fig