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During physiological stimulation by ACh or CCK, repetitive Ca 2+ spikes in the apical cytosol trigger both zymogen granule exocytosis and mitochondrial activation, ensuring that ATP generation is tightly coupled to the energetic cost of enzyme secretion ( 2+ signaling with ATP supply

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To the extent that depleted cysteine stores are restored, the component of ATP depletion caused by the ATP-depleting futile 5-oxoproline cycle will be ameliorated